Eopustularia of the Eocene

From the collection of Christian Romanek.
These exceptional shells allow a glimpse at one of the oldest lineages
of cowries from the Eocene (Lutetian and Bartonian) of France.
The genus Eopustularia (also popular under the name
Subepona) is ancestral to Propustularia and Nesiocypraea,
which are still present in the deep water of the Caribbean and the Indo-Pacific.
The shells depicted here were also used by Dolin & Aguerre (2018) who presented a comprehensive revision
of the Eocene members of the genus.